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英语翻译During the discovery and settlement of what is now the U

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英语翻译
During the discovery and settlement of what is now the United States,the flags of different European nations were flown over the land,as signs of possession
为啥是what,不是which
英语翻译During the discovery and settlement of what is now the U
during 后面的是插入语.把settlement去掉可能会好理解点(因为discovery和settlement并列的).during the discovery of someplace(先用这个词代替一下),what 引导了一个名词行丛句,what is now be called US.
再问: 这句话是什么意思