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An artist who seeks fame is like a dog chasing his own tail

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An artist who seeks fame is like a dog chasing his own tail who,结尾怎么加who
An artist who seeks fame is like a dog chasing his own tail who,when he captures it ,does not know what else to do but to coninue chasing it .
文中第2个who为什么加结尾,who单词什么时候加结尾,属于什么语法.
does not know what else to do but to continue chasing it 中的but to 怎么翻译解释.
An artist who seeks fame is like a dog chasing his own tail
1.第二个who引导定语从句(整个when he captures it ,does not know what else to do but to coninue chasing it 都由第二个who引导),who后加逗号只是为了停顿,因为后面这个句子太长了,没什么语法.
2.but to意思是:只有